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Here you can see the markings allowed to minors, majors, and chieftains respectively.
The only known Brute from this clan to ascend the Covenant hierarchy is Tartarus. Though this is disputed, as he is of mixed clan ancestry.
They have been known to bring larger members of their clan into battle on occassion, usually releasing them from their shackles as a self-destructive last resort.
*based of the Reach concept art of Brute redesign: http://www.isaachannaford.com/2011/05/brute-re-design-for-halo-reach.html Just a little photoshopping fun :P